名青少年史恩琼斯（纳森菲力普斯饰演），出去玩越野机车，穿越夏威夷的青葱绿地，却意外目睹了知名的洛杉矶检察官，惨遭恶名昭彰又行踪捉摸不定的帮派老大艾迪奇姆所杀害。奇姆希望这一击可以终结检方，针对他贩毒、勒索诈财、中介雏妓和枪枝买卖所进行的调查，不料，出现了目击证人，坏了他的好事。
现在，史恩是唯一亲眼看到他，犯下这桩狡猾残忍命案的人，只有史恩才能协助警方将他绳之以法。为了达到这个目的，首先必须说服史恩，站在洛杉矶大陪审团的面前出庭作证，于是要他离开夏威夷岛，前往美国本土，又不至于被奇姆派来的人灭口。这可不是一件简单的事，因为奇姆和他的手下，都很清楚史恩看见了什么，也知道史恩是检方瓦解他们犯罪帝国的唯一机会。他们会不计一切代价，拼了命地阻止他。
联邦调查局的探员奈佛弗林（山缪杰克森饰演），是处理危险状况的专家，他已经在史恩目击命案之后的几个小时，在艾迪奇姆的第一次刺杀行动中，救了史恩的一条命。弗林准备好要接受挑战，护送史恩前往洛杉矶，逮捕美国头号通缉要犯入狱。他的计划是利用一架私人飞机当成诱饵，同时征用一架商用客机的上层头等舱，偷偷将史恩带往洛杉矶，及时赶上出庭作证的时间。然而，有个机场的网民把消息泄露给奇姆一帮人，告诉他们护送史恩到洛杉矶的，是一架太平洋航空的子夜班机，而不是作为诱饵的私人喷射机，所以奇姆现在只要确定这架飞机到不了洛杉矶就好了，或者是让史恩永远无法活着下飞机。他要怎么办到呢？那就是在飞机上，偷偷放出能通过最严格安检的冷血动物……蛇。
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